'aggressive gray head,' photo from Ribbink et al. (1983)

A male Pseudotropheus "aggressive gray head" photographed at Maleri Island. This is another member of the informally recognized Pseudotropheus "aggressive" species group, and is different from the similarly named P. "aggressive gray". In the "aggressive gray head", Ribbink and colleagues found that males and females both defend territories, but that only about 60% of males, and no females, had territories in which algal "gardens" developed. The females allowed some mbuna of other species into their territories, which presumably managed to graze enough algae there to preclude the development of a "garden." Photo from Plate 7j of Ribbink et al., 1983; reproduced by permission of the Zoological Society of Southern Africa.

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